Contact Information:

Kathleen Pryer
Department of Biology
Duke University
Durham, NC 27707-0338
Telephone: 919-660-7380

Refereed Publications


Hirai, R.Y., E. Schuettpelz, L. Huiet, K.M. Pryer, A.R. Smith, and J. Prado. 2016. Phylogeny and relationships of the neotropical Adiantum raddianum group (Pteridaceae). Taxon 65: 1225–1235. Reprint PDF
Li, F.-W., L.-Y. Kuo, K.M. Pryer, C.J. Rothfels. 2016. Genes translocated into the plastid inverted repeat show decelerated substitution rates and elevated GC content. Genome Biology and Evolution 8: 2452–2458. Reprint PDF
Haufler, C.H., K.M. Pryer, E. Schuettpelz, E.B. Sessa, D.R. Farrar, R. Moran, J.J. Schneller, J.E. Watkins, Jr., and M.D. Windham. 2016. Sex and the single gametophyte: Revising the homosporous vascular plant life cycle in light of contemporary research. BioScience 66: 928-937. Reprint PDF
Rothfels, C.J., K.M. Pryer, F.-W. Li. 2016. Next-generation polyploid phylogenetics: Rapid resolution of hybrid polyploid complexes using PacBio single-molecule sequencing. New Phytologist. doi: 10.1111/nph.14111. Reprint PDF
Schuettpelz, E., C.-W. Chen, M. Kessler, J.B. Pinson, G. Johnson, A. Davila, A.T. Cochran, L. Huiet, and K.M. Pryer. 2016. A revised generic classification of vittarioid ferns (Pteridaceae) based on molecular, micromorphological, and geographic data. Taxon 65: 708–722. Reprint PDF
Pryer, K.M., L. Huiet, F.-W. Li, C.J. Rothfels, and E. Schuettpelz. 2016. Maidenhair ferns, Adiantum, are indeed monophyletic and sister to shoestring ferns, vittarioids (Pteridaceae). Systematic Botany 41: 17-23. Reprint PDF


2011 - 2015

Kao, T.-T., K.M. Pryer, M.D. Turner, R.A. White, and P. Korall. 2015. Origins of the endemic scaly tree ferns on the Galápagos and Cocos Islands. International Journal of Plant Sciences 176: 869-879.  Reprint PDF
Grusz, A.L. and K.M. Pryer. 2015. Development of microsatellite markers for the apomictic triploid fern, Myriopteris lindheimeri (Pteridaceae). Applications in Plant Sciences 3doi:10.3732/apps.1500061   Reprint PDF
Schuettpelz, E., K.M. Pryer, and M.D. Windham. 2015. A unified approach to taxonomic delimitation in the fern genus Pentagramma (Pteridaceae). Systematic Botany 40: 629-644. Reprint PDF
Wolf, P.G., E.B. Sessa, D.B. Marchant, F.-W. Li, C.J. Rothfels, E.M. Sigel, M.A. Gitzendanner, C.J. Visger, J.A. Banks, D.E. Soltis, P.S. Soltis, K.M. Pryer, and J.P. Der.  2015. An exploration into fern genome space. Genome Biology and Evolution 7: 2533-2544. Reprint PDF
Li, F.-W., C.J. Rothfels, M. Melkonian, J.C. Villarreal, D.W. Stevenson, S.W. Graham, G.K.-S. Wong, S. Mathews, and K.M. Pryer. 2015. The origin and evolution of phototropins. Frontiers in Plant Science 6: 637. Reprint PDF
Li, F.-W., M. Melkonian, C.J. Rothfels, J.C. Villareal, D.W. Stevenson, S.W. Graham, G.K.-S. Wong, K.M. Pryer, and S. Mathews.  2015Phytochrome diversity in green plants and the origin of canonical plant phytochromes. Nature Communications 6: 7852. Reprint PDF 
     MEDIA COVERAGE: Faculty 1000DukeTodayScience Daily,, NSF Science360 News
Rothfels C.J., F.-W. Li, E.M Sigel., L. Huiet, A. Larsson, D.O. Burge, M. Ruhsam, M. Deyholos, D. Soltis, N. Stewart, S. Shaw, L.M. Pokorny, T. Chen, C. dePamphilis, L. DeGironimo, D.W. Stevenson, S.W. Graham, G.K.-S. Wong, and K.M. Pryer. 2015The evolutionary history of ferns inferred from 25 low-copy nuclear genes. American Journal of Botany 102: 1089-1107. Reprint PDF
Huiet, L., M. Lenz, J.K. Nelson, K.M. Pryer, and A.R. Smith. 2015. Adiantum shastense, a new species of maidenhair fern from California.  Phytokeys 53: 73-81. Reprint PDF
Rothfels C.J., A.K. Johnson, P.H. Hovenkamp, D.L. Swofford, H.C. Roskam, C.R. Fraser-Jenkins, M.D. Windham, and K.M. Pryer. 2015. Natural hybridization between genera that diverged from each other approximately 60 million years ago. American Naturalist 185: 433-442. Reprint PDF
     MEDIA COVERAGENatureNPRAmerican,  Science Daily
Rothfels C.J., A.K. Johnson, M.D. Windham, and K.M. Pryer. 2014. Low-copy nuclear data confirm rampant allopolyploidy in the Cystopteridaceae (Polypodiales). Taxon 63: 1026-1036. Reprint PDF
Li, F.-W. and K.M. Pryer. 2014. Crowdfunding the Azolla fern genome project: a grassroots approach. GigaScience 3: 16.  Reprint PDF
Sessa, E.B., J.A. Banks, M.S. Barker, J.P. Der, A.M. Duffy, S.W. Graham, M. Hasebe, J. Langdale, F.-W. Li, D.B. Marchant, K.M. Pryer, C.J. Rothfels, S.J. Roux, M.L. Salmi, E.M. Sigel, D.E. Soltis, P.S. Soltis, D.W. Stevenson, and P.G. Wolf. 2014. Between two fern genomes. GigaScience 3: 15.  Reprint PDF
Pryer, K.M. 2014. "This lowly fern has massive green potential". The Toronto Globe and Mail 30 May 2014; "Offering an ancient solution to a modern problem". Contra Costa Times 1 June 2014. [Published Op-Ed].
Sigel, E.M., M.D. Windham, and K.M. Pryer. 2014. Evidence for reciprocal origins in Polypodium hesperium (Polypodiaceae): A natural fern model system for investigating how multiple origins shape allopolyploid genomes. American Journal of Botany 101: 1476–1485. Reprint PDF
Li, F.-W., J.C. Villarreal, S. Kelly, C.J. Rothfels, M. Melkonian, E. Frangedakis, M. Ruhsam, E.M. Sigel, J.P. Der, J. Pittermann, D.O. Burge, L. Pokorny, A. Larsson, T. Chen, S. Weststrand, P. Thomas, E. Carpenter, Y. Zhang, Z. Tian, L. Chen, Z. Yan, Y. Zhu, X. Sun, J. Wang, D.W. Stevenson, B.J. Crandall-Stotler, A.J. Shaw, M.K. Deyholos, D.E. Soltis, S.W. Graham, M.D. Windham, J.A. Langdale, G. K.-S. Wong, S. Mathews, and K.M. Pryer. 2014. Horizontal transfer of an adaptive chimeric photoreceptor from bryophytes to ferns. Proceedings of the National Academy of Sciences, USA 111: 6672-6677. Reprint PDF
    MEDIA COVERAGENew York TimesThe Economist, Current BiologyNational Geographic, Cosmos MagazineDuke Today, Scientific American, Decoded ScienceNature Reviews Genetics
Sigel, E.M., M.D. Windham, C.H. Haufler, and K.M. Pryer. 2014. Phylogeny, divergence time estimates, and phylogeography of the diploid species of the Polypodium vulgare complex. Systematic Botany 39: 1042-1055. Reprint PDF
Grusz, A.L., M.D. Windham, G. Yatskievych, L. Huiet, G.J. Gastony, and K.M. Pryer. 2014. Patterns of diversification in the xeric-adapted fern genus Myriopteris (Pteridaceae). Systematic Botany 39: 698-714, Reprint PDF.
Sigel, E.M., M.D. Windham, A.R. Smith, R.J. Dyer, and K.M. Pryer. 2014. Rediscovery of Polypodium calirhiza (Polypodiaceae) in Mexico. Brittonia 66: 278-286. Reprint PDF
Korall, P. and K.M. Pryer. 2014. Global biogeography of scaly tree ferns (Cyatheaceae): evidence for Gondwanan vicariance and limited transoceanic dispersal. Journal of Biogeography 41: 402-413. Reprint PDF
Rothfels C.J., A. Larsson, F.-W. Li, E.M. Sigel, L. Huiet, D.O. Burge, M. Ruhsam, S.W. Graham, D. Stevenson, G.K.-S. Wong, P. Korall, and K.M. Pryer. 2013. Transcriptome-mining for single-copy nuclear markers in ferns. PLoS ONE 8(10): e76957. doi:10:1371/journal.pone.0076957.  Reprint PDF
León, B., C.J. Rothfels, M. Arakaki, K.R. Young, and K.M. Pryer. 2013.  Revealing a cryptic fern distribution through DNA sequencing: Pityrogramma trifoliata in the western Andes of Peru. American Fern Journal 103. 40-48. Reprint PDF
Wang, Z.R. and K.M. Pryer. 2013. Gymnocarpium. Pp. 257-259 in Z.Y. Wu, P.H. Raven & D.Y. Hong, eds., Flora of China, Vol. 2–3 (Pteridophytes). Beijing: Science Press; St. Louis: Missouri Botanical Garden Press. Reprint PDF
Wang, Z.R., C.H. Haufler, K.M. Pryer, and M. Kato. 2013. Cystopteridaceae. Pp. 257 in Z.Y. Wu, P.H. Raven & D.Y. Hong, eds., Flora of China, Vol. 2–3 (Pteridophytes). Beijing: Science Press; St. Louis: Missouri Botanical Garden Press. Reprint PDF
Rothfels, C.J., M.D. Windham, and K.M. Pryer. 2013. A plastid phylogeny of the cosmopolitan fern family Cystopteridaceae (Polypodiopsida). Systematic Botany 38(2): 295–306. Reprint PDF
Pryer, K.M. 2012. The distinguished legacy of DMB: Donald MacPhail Britton (1923-2012). American Fern Journal 102: 241-251. Reprint PDF 
Beck, J.B., J. Allison, K.M. Pryer, and M.D. Windham. 2012. Identifying multiple origins of polyploid taxa: a multilocus study of the hybrid cloak fern (Astrolepis integerrima; Pteridaceae). American Journal of Botany 99: 1857-1865. Reprint PDF
Li, F.-W., K.M. Pryer, and M.D. Windham. 2012. Gaga, a new fern genus segregated from Cheilanthes (Pteridaceae). Systematic Botany 37: 845-860. Reprint PDF
  MEDIA COVERAGE: Rolling StoneWiredHuffington PostNational GeographicGuardianNew York Times
Johnson, A.K., C.J. Rothfels, M.D. Windham, and K.M. Pryer. 2012. Unique expression of a sporophytic character on the gametophytes of notholaenid ferns (Pteridaceae). American Journal of Botany 99: 1118-1124. Reprint PDF
Rothfels, C.J.*, M.A. Sundue*, L.-Y. Kuo, A. Larsson, M. Kato, E. Schuettpelz, and K.M. Pryer. 2012. A revised family-level classification for eupolypod II ferns (Polypodiidae: Polypodiales). Taxon 61: 515-533. *equally contributing. Reprint PDF
Rothfels, C.J.*, A. Larsson*, L.-Y. Kuo*, P. Korall, W.-L. Chiou, and K.M. Pryer. 2012. Overcoming deep roots, fast rates, and short internodes to resolve the ancient rapid radiation of eupolypod II ferns. Systematic Biology 61: 490-509. doi: 10.1093/sysbio/sys001. *equally contributing. Reprint PDF (Cover)
Li, F.-W., L.-Y. Kuo, C.J. Rothfels, A. Ebihara, W.-L. Chiou, M.D. Windham, and K.M. Pryer. 2011. rbcL and matK earn two thumbs up as the core DNA barcode for ferns. PLoS One 6: e26597. Reprint PDF
Beck, J.B., M.D. Windham, and K.M. Pryer. 2011. Do asexual lineages lead short evolutionary lives? A case-study from the fern genus Astrolepis. Evolution 65: 3217-3229. Reprint PDF 
Sigel, E.M., M.D. Windham, L. Huiet, G. Yatskievych, and K.M. Pryer. 2011. Species relationships and farina evolution in the cheilanthoid fern genus Argyrochosma (Pteridaceae). Systematic Botany 36: 554-564. Reprint PDF
Wolf, P.G., J.P. Der, A.M. Duffy, J.B. Davidson, A.L. Grusz, and K.M. Pryer. 2011. The evolution of chloroplast genes and genomes in ferns. Plant Molecular Biology 76: 251–261. Reprint PDF


2000 - 2010

Pryer, K.M., E. Schuettpelz, L. Huiet, A.L. Grusz, C.J. Rothfels, T. Avent, D. Schwartz, and M.D. Windham. 2010. DNA barcoding exposes a case of mistaken identity in the fern horticultural trade. Molecular Ecology Resources 10: 979-985. Reprint PDF
Korall, P., E. Schuettpelz, and K.M. Pryer. 2010. Abrupt deceleration of molecular evolution linked to the origin of arborescence in ferns. Evolution 64: 2786-2792. Reprint PDF
Beck, J.B., M.D. Windham, G. Yatskievych, and K.M. Pryer. 2010. A diploids-first approach to species delimitation and interpreting polyploid evolution in the fern genus Astrolepis (Pteridaceae). Systematic Botany 35: 223-234. Reprint PDF
Schneider, H., A.R. Smith, and K.M. Pryer. 2009. Is morphology really at odds with molecules in estimating fern phylogeny? Systematic Botany 34: 455-475. Reprint PDF
Windham, M.D., L. Huiet, E. Schuettpelz, A.L. Grusz, C. Rothfels, J. Beck, G. Yatskievych, and K.M. Pryer. 2009. Using plastid and nuclear DNA sequences to redraw generic boundaries and demystify species complexes in cheilanthoid ferns. American Fern Journal 99: 128-132. Reprint PDF
Grusz, A.L., M.D. Windham, and K.M. Pryer. 2009. Deciphering the origins of apomictic polyploids in the Cheilanthes yavapensis complex (Pteridaceae). American Journal of Botany 96: 1636-1645. Reprint PDF
Schuettpelz, E. and K.M. Pryer. 2009. Evidence for a Cenozoic radiation of ferns in an angiosperm-dominated canopy. Proceedings of the National Academy of Sciences, USA 106: 11200-11205 (cover). Reprint PDF  Cover
Pryer, K.M. and D.J. Hearn. 2009. Evolution of leaf form in marsileaceous ferns: evidence for heterochrony. Evolution 63: 498-513. Reprint PDF
Pryer, K.M. and E. Schuettpelz. 2009. Ferns. Chapter 14 in S.B. Hedges and S. Kumar (eds.) Timetree of Life. Oxford University Press, New York. pp. 153-156.  Reprint PDF
Schuettpelz, E., A.L. Grusz, M.D. Windham, and K.M. Pryer. 2008. The utility of nuclear gapCp in resolving polyploid fern origins. Systematic Botany 33: 621-629. Reprint PDF
Hennequin, S., E. Schuettpelz, K.M. Pryer, A. Ebihara, and J.-Y. Dubuisson. 2008. Divergence times and the evolution of epiphytism in filmy ferns (Hymenophyllaceae) revisited. International Journal of Plant Sciences 169: 1278-1287. Reprint PDF
Nagalingum, N.S., M.D. Nowak, and K.M. Pryer. 2008. Assessing phylogenetic relationships in extant heterosporous ferns (Salviniales), with a focus on Pilularia and Salvinia. Botanical Journal of the Linnaean Society 157: 673-685. Reprint PDF
Rothfels, C.J., M.D. Windham, A.L. Grusz, G.J. Gastony, and K.M. Pryer. 2008. Toward a monophyletic Notholaena (Pteridaceae): Resolving patterns of evolutionary convergence in xeric-adapted ferns. Taxon 57: 712-724. Reprint PDF
Schuettpelz, E. and K.M. Pryer. 2008. Fern Phylogeny. Chapter 15 in Ranker, T.A. and C.H. Haufler (eds.). The Biology and Evolution of Ferns and Lycophytes. Cambridge Univ. Press., pp. 395-416. Reprint PDF
Smith A.R., K.M. Pryer, E. Schuettpelz, P. Korall, H. Schneider, and P.G. Wolf. 2008. Fern Classification. Chapter 16 in Ranker, T.A. and C.H. Haufler (eds.). The Biology and Evolution of Ferns and Lycophytes. Cambridge Univ. Press., pp. 417-467. Reprint PDF
Metzgar, J.S., J.E. Skog, E.A. Zimmer, and K.M. Pryer. 2008. The paraphyly of Osmunda is confirmed by phylogenetic analyses of seven plastid loci. Systematic Botany 33: 31-36. Reprint PDF
Christenhusz, M.J.M., H. Tuomisto, J.S. Metzgar, and K.M. Pryer. 2008. Evolutionary relationships within the Neotropical, eusporangiate fern genus Danaea (Marattiaceae). Molecular Phylogenetics and Evolution 46: 34-48. Reprint PDF
Schuettpelz E. and K.M. Pryer. 2007. Fern phylogeny inferred from 400 leptosporangiate species and three plastid genes. Taxon 56: 1037-1050. Reprint PDF
Haugen, P., D. Bhattacharya, J.D. Palmer, S. Turner, L.A. Lewis, and K.M. Pryer. 2007. Cyanobacterial ribosomal RNA genes with multiple, endonuclease-encoding group I introns. BMC Evolutionary Biology 7: 159. Reprint PDF
Metzgar, J.S., H. Schneider, and K.M. Pryer. 2007. Phylogeny and divergence time estimates for the fern genus Azolla (Salviniaceae). International Journal of Plant Sciences 168: 1045-1053. Reprint PDF
Schuettpelz E., H. Schneider, L. Huiet, M.D. Windham, and K.M. Pryer. 2007. A molecular phylogeny of the fern family Pteridaceae: assessing overall relationships and the affinities of previously unsampled genera. Molecular Phylogenetics and Evolution 44: 1172-1185. Reprint PDF
Korall, P., D.S. Conant, J.S. Metzgar, H. Schneider, and K.M. Pryer. 2007. A molecular phylogeny of scaly tree ferns (Cyatheaceae). American Journal of Botany 94: 873-886 (cover). Reprint PDF
Nagalingum, N.S., H. Schneider, and K.M. Pryer. 2007. Molecular phylogenetic relationships and morphological evolution in the heterosporous fern genus Marsilea. Systematic Botany 32: 16-25. Reprint PDF
Schuettpelz, E., P. Korall, and K.M. Pryer. 2006. Plastid atpA data provide improved support for deep relationships among ferns. Taxon 55: 897-906. Reprint PDF
Schuettpelz, E. and K.M. Pryer. 2006. Reconciling extreme branch length differences: decoupling time and rate through the evolutionary history of filmy ferns. Systematic Biology 55: 485-502. [2006 Publisher's Award for Excellence in Systematic Research]. Reprint PDF
Korall, P., K.M. Pryer, J. Metzgar, H. Schneider, and D.S. Conant. 2006. Tree ferns: monophyletic groups and their relationships as revealed by four protein-coding plastid loci. Molecular Phylogenetics and Evolution 39: 830-845. Reprint PDF
Nagalingum, N.S., H. Schneider, and K.M. Pryer. 2006. Comparative morphology of reproductive structures in heterosporous water ferns and a reevaluation of the sporocarp. International Journal of Plant Sciences 167: 805-815. Reprint PDF
Korall, P., D.S. Conant, H. Schneider, K. Ueda, H. Nishida, and K.M. Pryer. 2006. On the phylogenetic position of Cystodium: it’s not a tree fern – it's a polypod! American Fern Journal 96: 45-53. Reprint PDF
Smith, A.R., K.M. Pryer, E. Schuettpelz, P. Korall, H. Schneider, and P.G. Wolf. 2006. A classification for extant ferns. Taxon 55: 705-731. Reprint PDF    For translation into Ukrainian click here
Wikström, N. and K.M. Pryer. 2005. Incongruence between primary sequence data and the distribution of a mitochondrial atp1 group II intron among ferns and horsetails. Molecular Phylogenetics and Evolution 36: 484-493. Reprint PDF
Schneider, H., E. Schuettpelz, K.M. Pryer, R. Cranfill, S. Magallón, R. Lupia. 2004. Ferns diversified in the shadow of angiosperms. Nature 428: 553-557. Reprint PDF
Pryer, K.M., H. Schneider and S. Magallón. 2004. The radiation of vascular plants. In J. Cracraft and M. J. Donoghue, eds. Assembling the Tree of Life, 138-153. Oxford University Press, London. Reprint PDF
Pryer, K.M., E. Schuettpelz, P.G. Wolf, H. Schneider, A.R. Smith, R. Cranfill. 2004. Phylogeny and evolution of ferns (monilophytes) with a focus on the early leptosporangiate divergences. American Journal of Botany 91: 1582-1598. [Special invited paper]  Reprint PDF
Des Marais, D.L., A.R. Smith, D.M. Britton, and K.M. Pryer. 2003. Phylogenetic relationships and evolution of extant horsetails,Equisetum, based on chloroplast DNA sequence data (rbcL andtrnL-F). International Journal of Plant Sciences 164: 737-751. Reprint PDF
Dubuisson, J.-Y., S. Hennequin, E.J.P. Douzery, R. Cranfill, A.R. Smith and K.M. Pryer. 2003. rbcL phylogeny of the fern genusTrichomanes (Hymenophyllaceae), with special reference to neotropical taxa. International Journal of Plant Sciences 164: 753-761. Reprint PDF
Pryer, K.M., H. Schneider, E.A. Zimmer and J.A. Banks. 2002. Deciding among green plants for whole genome studies. Trends in Plant Science 7: 550-554. Reprint PDF
Schneider, H. and K.M. Pryer. 2002. Structure and function of spores in the aquatic heterosporous fern family Marsileaceae. International Journal of Plant Sciences 163: 485-505 (cover). Reprint PDF
Schneider, H., K.M. Pryer, R. Cranfill, A.R. Smith and P.G. Wolf. 2002. Evolution of vascular plant body plans: a phylogenetic perspective. Pp. 330-364 in Developmental Genetics and Plant Evolution, Q.C.B. Cronk, R.M. Bateman, and J.A. Harris, eds. Taylor and Francis, Philadelphia. Reprint PDF
Smith, A.R., H. Tuomisto, K.M. Pryer, J.S. Hunt and P.G. Wolf. 2001. Metaxya lanosa, a second species in the genus and fern family Metaxyaceae. Systematic Botany 26: 480-486. Reprint PDF
Pryer, K.M., A.R. Smith, J.S. Hunt, and J.-Y.Dubuisson. 2001. rbcLdata reveal two monophyletic groups of filmy ferns (Filicopsida:Hymenophyllaceae). American Journal of Botany 88: 1118-1130 (cover). Reprint PDF
Pryer, K.M., H. Schneider, A.R. Smith, R. Cranfill, P.G. Wolf, J.S. Hunt and S.D. Sipes. 2001. Horsetails and ferns are a monophyletic group and the closest living relatives to seed plants. Nature 409: 618-622 (cover). Reprint PDF
Lupia, R., H. Schneider, G.M. Moeser, K.M. Pryer and P.R. Crane. 2000. Marsileaceae sporocarps and spores from the Late Cretaceous of Georgia, U.S.A. International Journal of Plant Sciences 161: 975-988. Reprint PDF


Prior to 2000

Wolf, P.G., S.D. Sipes, M.R. White, M.L. Martines, K.M. Pryer, A.R. Smith and K. Ueda. 1999. Phylogenetic relationships of the enigmatic fern families Hymenophyllopsidaceae and Lophosoriaceae: evidence from rbcL nucleotide sequences. Plant Systematics and Evolution 219: 263-270. Reprint PDF
Turner, S., K.M. Pryer, V.P.W. Miao and J.D. Palmer. 1999. Investigating deep phylogenetic relationships among cyanobacteria and plastids by small subunit rRNA sequence analysis. Journal of Eukaryotic Microbiology 46: 327-338. Reprint PDF
Pryer, K.M. 1999. Phylogeny of marsileaceous ferns and relationships of the fossil Hydropteris pinnata reconsidered. International Journal of Plant Sciences 160:931-954 (cover). Reprint PDF
Wolf, P.G., K.M. Pryer, A.R. Smith and M. Hasebe. 1998. Phylogenetic studies of extant pteridophytes. Pp. 541-556 in D.E. Soltis, P.S. Soltis and J.J. Doyle, eds. Molecular systematics of plants II: DNA sequencing. Kluwer Academic Publishers, Boston. Reprint PDF
Pryer, K.M., A.R. Smith and J.E. Skog. 1995. Phylogenetic relationships of extant ferns based on evidence from morphology and rbcL sequences. American Fern Journal 85 (Special Issue): 205-282. Reprint PDF
Hasebe, M., P.G. Wolf, K.M. Pryer, K. Ueda, M. Ito, R. Sano, G.J. Gastony, J. Yokoyama, J.R. Manhart, N. Murakami, E.H. Crane, C.H. Haufler and W.D. Hauk. 1995. Fern phylogeny based on rbcLnucleotide sequences. American Fern Journal 85 (Special Issue): 134-181. Reprint PDF
Pryer, K.M. and C.H. Haufler. 1993. Isozymic and chromosomal evidence for the allotetraploid origin of Gymnocarpium dryopteris(Dryopteridaceae). Systematic Botany 18: 150-172. Reprint PDF
Pryer, K.M. 1993. Gymnocarpium. Pp. 258-262 in Flora of North America North of Mexico, Vol.2. Flora of North America Editorial Committee (ed.). Oxford University Press, New York. Reprint PDF
Pryer, K.M. 1992. The status of Gymnocarpium heterosporum and G. robertianum in Pennsylvania. American Fern Journal 81: 34-39. Reprint PDF
Pryer, K.M. 1990. The limestone oak fern: new to Manitoba. The Blue Jay 48:192-195. Reprint PDF
Pryer, K.M. and L.R. Phillippe. 1989. A synopsis of the genus Sanicula in eastern Canada. Canadian Journal of Botany 67: 694-707. Reprint PDF
McNeill, J. and K.M. Pryer. 1985. The status and typification of Phegopteris and Gymnocarpium. Taxon 34: 136-156. Reprint PDF
Pryer, K.M., D.M. Britton and J. McNeill. 1983. A numerical analysis of chromatographic profiles in North American taxa of the fern genus Gymnocarpium. Canadian Journal of Botany 61: 2592-2602. Reprint PDF
Pryer, K.M. and D.M. Britton. 1983. Spore studies in the genus Gymnocarpium. Canadian Journal of Botany 61: 377-388. Reprint PDF 
Sarvela, J., D.M. Britton and K.M. Pryer. 1981. Studies on the Gymnocarpium robertianum complex in North America. Rhodora 83: 421-431. Reprint PDF
Chase, R., K.M. Pryer, R. Baker and D.M. Madison. 1978. Responses to conspecific chemical stimuli in the terrestrial snail Achatina fulica (Pulmonata: Sigmuretha). Behavioural Biology 22: 302-315. Reprint PDF



Argus, G.W. & K.M. Pryer. 1990. Rare Vascular Plants in Canada - Our Natural Heritage. Canadian Museum of Nature, Ottawa. 277 pp.
Pryer, K.M. & G.W. Argus (eds.). 1987. Atlas of the Rare Vascular Plants of Ontario. Part 4. National Museum of Natural Sciences, Ottawa. 472 pp.  


Electronic Publications

Pryer, Kathleen M., Alan R. Smith, and Carl Rothfels. 2008. Polypodiidae Cronquist, Takht. & Zimmerm. 1966. Leptosporangiate Ferns. Version 23 December 2008. in The Tree of Life Web Project,
Pryer, Kathleen M., Alan R. Smith, and Carl Rothfels. 2009. Polypodiopsida Cronquist, Takht. & Zimmerm. 1966. Ferns. Version 14 January 2009. in The Tree of Life Web Project,
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